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Donlevy's break came in 1935, when he was cast in the Edward G. Robinson film Barbary Coast.
A large amount of film work followed, with several important parts.
In 1939, he played the lead villain in Destry Rides Again and was nominated for an Academy Award for Best Supporting Actor for his role as the ruthless Sergeant Markoff in Beau Geste, although the Oscar went to Thomas Mitchell for Stagecoach.

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