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Grand Duke Dmitri Pavlovich was born at Ilinskoe near Moscow, the second child and son of Grand Duke Paul Alexandrovich and a grandson of Alexander II of Russia ; thus, he was a first cousin of Nicholas II of Russia.
Dmitri Pavlovich's mother, Alexandra Georgievna of Greece was a daughter of George I of Greece and his Queen consort, Olga Konstantinovna of Russia.
As such, he was also a first cousin of Prince Philip, Duke of Edinburgh.
Dmitri and his sister Maria were mostly raised by their uncle and aunt, Grand Duke Sergei Alexandrovich of Russia and his wife, Grand Duchess Elizabeth Fyodorovna, the elder sister of Tsarina Alexandra.

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