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In Vespa crabro, the nest is founded in spring by a fertilized female known as the queen.
She generally selects sheltered places like dark hollow tree trunks.
She first builds a series of cells ( up to 50 ) out of chewed tree bark.
The cells are arranged in horizontal layers named combs, each cell being vertical and closed at the top.
An egg is then laid in each cell.
After 5 – 8 days, the egg hatches, and in the next two weeks, the larva undergoes its five stages.
During this time, the queen feeds it a protein-rich diet of insects.
Then, the larva spins a silk cap over the cell's opening and, during the next two weeks, transforms into an adult, a process called metamorphosis.
The adult then eats its way through the silk cap.
This first generation of workers, invariably females, will now gradually undertake all the tasks formerly carried out by the queen ( foraging, nest building, taking care of the brood, etc.
) with one exception: egg-laying, which remains exclusive to the queen.

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