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The scribe, usually a monk, would decide on what his quire ( four folded sheets ) would look like, by arranging the hair and flesh sides of the sheets.
Throughout time from Carolingian period and all the way up to the Middles Ages, different styles of folding the quire came about.
For example, in mainland Europe throughout the Middle Ages, the quire would be put into a system which each side would fold on to the same style.
The hair side would meet the hair side and the same goes with the flesh side.
This was not the same style used in the British Isles, where the membrane would be folded so that it turned out an eight leaf quire, with single leaves in the third and sixth positions.
Once the scribe has it the way that he wants, the next stage was tacking the quire.
Tacking is when the scribe would hold together the leaves in quire with thread.
Once threaded together, the scribe would then sew a line of parchment up the “ spine ” of the manuscript, as to protect the tacking.

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