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A field of study called memetics arose in the 1990s to explore the concepts and transmission of memes in terms of an evolutionary model.
Criticism from a variety of fronts has challenged the notion that academic study can examine memes empirically.
However, developments in neuroimaging may make empirical study possible.
Some commentators question the idea that one can meaningfully categorize culture in terms of discrete units.
Others, including Dawkins himself, have argued that this usage of the term is the result of a misunderstanding of the original proposal.

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