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It is hypothesized that their survival was based on the benefits of very large numbers.
There was safety in large flocks which often numbered hundreds of thousands of birds.
When a flock of this huge a size established itself in an area, the number of local animal predators ( such as wolves, foxes, weasels, and hawks ) was so small compared to the total number of birds that little damage would be inflicted on the flock as a whole.
It was common for passenger pigeon flocks to perch on each other's backs, an unusual behavior even for socially inclined birds.

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