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The Pemalites were destroyed several thousand years before the main events of the series.
The race was targeted for extermination by the Howlers, almost surely because their master, Crayak, opposed their service to his enemy the Ellimist.
Having no weapons, the species was almost completely wiped out.
The Pemalites who survived the initial attacks abandoned their planet and, along with the Chee, went to Earth.
However, the Howlers had unleashed a genetic virus along with their attacks, and as a result the few remaining members of the species began to die, and reached Earth barely clinging to life.
The Chee managed to save their essence, however, and transferred this into wolves, resulting in the creation of the domesticated dog, which, in the words of Erek King, are " all that remains of the Pemalite soul.

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