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A third explanation is suggested by Richardson's analysis of Sukuma tone.

There we see a basically simple phonemic system enmeshed in a very complex and puzzling morphophonemic system.

While the phonemes can be very easily stated, no one is likely to be satisfied with the statement until phonemic occurrences can be related in some way to morphemic units, i.e. until the morphophonemics is worked out, or at least far enough that it seems reasonable to expect success.