** Rüppell's Horseshoe Bat, Rhinolophus fumigatus

**** Rüppell's Broad-nosed Bat, Scoteanax rueppellii

* Rüppell's Broad-Nosed Bat, Scoteanax rueppellii

* Andaman Horseshoe Bat ( Rhinolophus cognatus )

Small numbers of Greater Horseshoe Bat ( Rhinolophus ferrumequinum ), Lesser Horseshoe Bat ( Rhinolophus hipposideros ) and Natterer's bat ( Myotis nattereri ) hibernate in the cave system.

He also described the Lesser Horseshoe Bat for the first time.

** Adam's Horseshoe Bat, Rhinolophus adami

** Maendeleo Horseshoe Bat, Rhinolophus maendeleo

** Cape Horseshoe Bat, Rhinolophus capensis

** Dent's Horseshoe Bat, Rhinolophus denti

** Bushveld Horseshoe Bat, Rhinolophus simulator

** Swinny's Horseshoe Bat, Rhinolophus swinnyi

** Mediterranean Horseshoe Bat, Rhinolophus euryale

** Mehely's Horseshoe Bat, Rhinolophus mehelyi

** Arcuate Horseshoe Bat, Rhinolophus arcuatus ( see also Andersen's Horseshoe Bat )

** Canut's Horseshoe Bat, Rhinolophus canuti

** Croslet Horseshoe Bat, Rhinolophus coelophyllus

** Creagh's Horseshoe Bat, Rhinolophus creaghi

** Broad-eared Horseshoe Bat, Rhinolophus euryotis

** Philippine Forest Horseshoe Bat, Rhinolophus inops

** Large Rufous Horseshoe Bat, Rhinolophus rufus

** Shamel's Horseshoe Bat, Rhinolophus shameli

** Small Rufous Horseshoe Bat, Rhinolophus subrufus

** Bokhara Horseshoe Bat, Rhinolophus bocharicus

** Species: Alphacoronavirus 1, Human coronavirus 229E, Human Coronavirus NL63, Miniopterus Bat coronavirus 1, Miniopterus Bat coronavirus HKU8, Porcine epidemic diarrhea virus, Rhinolophus Bat coronavirus HKU2, Scotophilus Bat coronavirus 512