However, his relatives were not major court officials ; documents show that El Cid's paternal grandfather, Lain, confirmed only five documents of Ferdinand I's, his maternal grandfather, Rodrigo Alvarez, certified only two of Sancho II's, and El Cid's own father confirmed only one.

However, our understanding of the molecular mechanisms behind genomic imprinting show that it is the maternal genome that controls much of the imprinting of both its own and the paternally-derived genes in the zygote, making it difficult to explain why the maternal genes would willingly relinquish their dominance to that of the paternally-derived genes in light of the conflict hypothesis.

However, Brando was closer to his maternal grandmother, Bessie Gahan Pennebaker Meyers, than to his mother.

However, the country lagged behind on the goals of eliminating extreme poverty and hunger, reducing child and maternal mortality, and combating diseases such as human immunodeficiency virus / acquired immune deficiency syndrome ( HIV / AIDS ) and malaria.

However, their modes of maternal provisioning vary greatly.

However, by the time of the novel Pastime, Spenser states that his mother died during labor and he was delivered via Caesarian section, i. e. " not of woman born " as Parker has Spenser put it ; he was raised by his father and his two maternal uncles.

However, preterm births can suffer neurodevelopmental disorders due to lack of maternal thyroid hormones due their own thyroid being insufficiently developed to meet their postnatal needs.

However, that study was later found to be based on incomplete data, and recent studies point to multiple maternal origins, with the clade found in the Americas, Europe, Middle East, and Africa, originating from the Indian subcontinent, where a large number of unique haplotypes occur.

However her mother, Madeleine de la Tour d ' Auvergne, the Countess of Boulogne, was from one of the most prominent and ancient French noble families ; this prestigious maternal heritage was of benefit to her future marriage to a Royal Prince of France.

However, the Dictionary of the Second Empire states that Haussmann used the title of baron casually, out of pride as the only male descendant of his maternal grandfather, Georges Frédéric, Baron Dentzel, a general under the first Napoleon.

However, both feature films and the animated television series conformed to Charles Addams's original concept of Grandmama as Wednesday's and Pugsley's maternal grandmother.

However, he turned out to be a noted broodmare sire, being the maternal grandsire (" damsire ") of 1992 Horse of the Year and successful sire A. P.

However, an argument began as to who should take the boar's skin as a prise: Meleager gave it to Atalanta, but two of his maternal uncles, sons of Thestius, wanted the trophy for themselves, claiming that it belonged to them by the right of birth if Meleager did not want it.

However, John was the eldest ( although maternal ) grandson of the 1st Duke of the 1st creation.

However, not wishing to repeat the difficulties of three decades previous, Philip, a few months before his marriage, renounced his princely titles and adopted the surname Mountbatten, which was that of his uncle and mentor, the Viscount Mountbatten of Burma, and itself was adopted by the Viscount's father ( Philip's maternal grandfather ), Prince Louis of Battenberg, in 1917.

However, inheritance of wealth does not follow the patriclan but is determined by the matriclan, that is, a son does not inherit his father's cattle but his maternal uncle's instead.

However, Kurdish groups show a closer relationship with European groups than with Caucasian groups based on mtDNA, but the opposite based on the Y-chromosome, indicating some differences in their maternal and paternal histories.

However, some AVP may also be released directly into the brain, and accumulating evidence suggests it plays an important role in social behavior, bonding, and maternal responses to stress.

However, it is notable that during much of that film, Giselle was more of an older sister figure than a maternal figure to that little girl.

However, there appears to be no correlation between HPE and maternal age.

However, testing the mitochondrial genome can prove only if two individuals are related by common descent through maternal lines only from a common ancestor and is, thus, of limited value ( for instance, it could not be used to test for paternity ).

However, there have been several cases where a descendant through the maternal line has changed their surname in order to claim the chiefship of a clan, such as the late chief of the Clan MacLeod who was born John Wolridge-Gordon and changed his name to the maiden name of his maternal grandmother in order to claim the chiefship of the MacLeods.

However, just as all the buildings have not fallen and flowed back to their original mud, so the values which wanted them and saw that they were built have not all disappeared.

However, Biblical scholars frequently attested to its numerous inaccuracies, as old manuscripts were uncovered and scholarship advanced.

However, last night the tapes were not run at all during the evening hours and all we got on TV were a few snatches which Douglas Edwards and Huntley and Brinkley could squeeze into their programs.

However, it determined that neither this factor, nor `` the fact that all concerned in high executive posts in both companies acted honorably and fairly, each in the honest conviction that his actions were in the best interests of his own company and without any design to overreach anyone, including Du Pont's competitors '', outweighed the Government's claim for relief.

However, they do demonstrate the presence of large normal pressures in the presence of flat shear fields which were forecast by the theory in the first part of the paper.

However, the statistical offices of both governments were assigned responsibility for the planning and analysis of these statistics.

However, they were hostile to Jewish Orthodoxy, professing to believe in Judaism `` but in a moderate way ''.

However, the positions of all parties and leaders were constantly shifting.

However, the wei books were also destroyed in a series of Orthodox Confucian purges which culminated in a final proscription in 605.

However, while usually Greek festivals were celebrated at the full moon, all the feasts of Apollo were celebrated at the seventh day of the month, and the emphasis given to that day ( sibutu ) indicates a Babylonian origin.

However, recent work suggests that some agnostids were benthic predators, engaging in cannibalism and possibly manifesting pack-hunting behavior.

However, they were generally occupied for a mere 30 years or less.

However, data from the National Statistical Service shows that there were only 65, 000 foreigners staying in Armenian hotels in 2009.

However, by mid-1837 the South Australian Register was warning of escaped convicts from New South Wales, and tenders for a temporary gaol were sought.

However, these claims were given up in about 1870 during the War of the Pacific between Chile, the allied Bolivia and Peru, in a diplomatic deal to keep Argentina out of the war.

However, tyrannosaurid forelimbs were extremely small for their body size and retained only two digits.

However, this theory is undermined by the disproportional fear of spiders in comparison to other, potentially dangerous creatures that were present during Homo sapiens environment of evolutionary adaptiveness.

However, no incidents of violence against spring breakers were reported.

However, if we look for the peoples in the region from the upper Main in the north, south to the Danube and east to the Czech Republic where the Quadi and Marcomanni were located, Ptolemy does not give any tribes.

However, this was halted as a report reached Constantinople that the Safavids were invading the Ottoman Empire, causing a period of panic, turning the Sultan's attention away from Russia.

However, Margaret Sleeboom-Faulkner of the Department of Anthropology at the University of Sussex said Kanzō Umehara considered the Ainu and some Ryukyuans to have " preserved their proto-Mongoloid traits " According to anthropologist Elsie Clews Parsons physical features of the Proto-Mongoloid were characterized as, " a straight-haired type, medium in complexion, jaw protrusion, nose-breadth, and inclining probably to round-headedness ".

However, before they were to gain entry into the sanctuary, Straboromanos and royal guards caught up with them to summon them back to the palace.

However, later that month, on 22 January, the English were defeated at the Battle of Basing and, on the 22 March at the Battle of Merton ( perhaps Marden in Wiltshire or Martin in Dorset ), in which Æthelred was killed.