They were split based on pronounced differences in morphology and anatomy ; two molecular studies, each based on a very small number of genes, indicate that the species are distinct but disagree over how closely related they are.

They were split based on pronounced differences in morphology and anatomy ( Clark, 1992 ; Olson, 1994 ; Sangsteret al., 2002 ); two molecular studies, each based on a very small number of genes, indicate that the species are distinct but disagree over how closely related they are.

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