After centriole duplication, the two pairs of centrioles remain attached to each other in an orthogonal configuration until mitosis, when the mother and daughter centrioles separate in a manner dependent upon the enzyme separase.

After the First World War, the railway companies were rationalised and merged into the Southern Railway which set about removing duplication from the railway system.

After the R-point has been passed in “ G1b ”, however, extracellular signals are no longer required, and the cell is irreversibly committed to preparing for DNA duplication.

After the end of apartheid allowed the consideration of the entire city of Johannesburg as one without consideration of race, it was determined that the previous structure of the city was wasteful and that there was much duplication of functions.

< gallery caption =" After duplication ">

After 1890 came philosopher Josiah Royce ( 1855 – 1916 ), botanist Liberty Hyde Bailey ( 1858 – 1954 ), the Southern Agrarians of the 1920s and 1930s, novelist John Steinbeck ( 1902 – 1968 ), historian A. Whitney Griswold ( 1906 – 1963 ), environmentalist Aldo Leopold ( 1887 – 1948 ), Ralph Borsodi ( 1886 – 1977 ), and present-day authors Wendell Berry ( b. 1934 ), Gene Logsdon ( b. 1932 ), Paul Thompson, and Allan C. Carlson ( b. 1949 ).

After tensions with the U. S. government came to a head in 1890, the church officially abandoned the public practice of polygamy in the United States, and eventually stopped performing official polygamous marriages altogether after a Second Manifesto in 1904.

After that, censuses followed somewhat regularly in 1787, 1801, and 1834, and between 1840 and 1860, the censuses were taken every five years, and then every ten years until 1890.

After the removal of the powerful Chancellor Otto von Bismarck in 1890 following the death of Emperor Wilhelm I, the young Emperor Wilhelm II engaged in increasingly reckless foreign policies that left the Empire isolated.

After Germany was united by Otto von Bismarck into the " Second German Reich ", he determined German politics until 1890.

After his father's death, Edmund Gosse published a typical Victorian biography, The Life of Philip Henry Gosse ( 1890 ).

After graduating from Perm gymnasium in 1890, he went to the capital to study law at St. Petersburg University, but ended up also taking classes at the St. Petersburg Conservatory of Music, where he studied singing and music ( a love of which he had picked up from his stepmother ).

After some years spent teaching and in ineffective attempts to obtain the production of more than one opera, he saw the enormous success of Pietro Mascagni's Cavalleria rusticana in 1890, and he wasted no time in producing his own verismo hit, Pagliacci.

After being ratified in the Senate on April 8, 1890 by a vote of 51-1, the Sherman Act passed unanimously ( 242-0 ) in the House of Representatives on June 20, 1890, and was then signed into law by President Benjamin Harrison on July 2, 1890.

After immense hardships and great loss of life, Stanley met Emin in 1888, charted the Ruwenzori Range and Lake Edward, and emerged from the interior with Emin and his surviving followers at the end of 1890.

After a brief de facto protectorate period beginning in 1885 the island became a full formal French protectorate in 1890, then a colony in 1896, and gained full independence from France in 1960 in the wake of decolonization.

After 450 amendments, the Tariff Act of 1890 was passed, and increased average duties across all imports from 38 % to 49. 5 %.

After only six months ' work he completed his first picture, which was exhibited at the Paris Salon of 1890.

After 1890 Strauss composed very infrequently for chamber groups, his energies being almost completely absorbed with large-scale orchestral works and operas.

After an August 5, 1890 election, the voters of County 5 overwhelmingly voted for the name " Kingfisher " over " Hennesey " and " Harrison "

After the passage of the Organic Act in 1890, Cleveland County was organized as County 3 and Norman became the county seat.

After becoming part of the Oklahoma Territory in 1890, Beaver County ( first called Seventh County ) covered the entire Oklahoma Panhandle.

After Idaho statehood in 1890, as in the rest of the state, mining gradually decreased in significance in Blaine County.

After the Holy Trinity Church in Leland burned down in 1880, the first wood-frame church was built in 1890 and the cemetery was established.

After becoming part of the newly formed Atlantic County in 1837, portions of the township were taken to create Mullica Township ( March 13, 1838 ), Egg Harbor City ( June 14, 1858 ), Absecon town ( February 29, 1872 ), Brigantine Beach borough ( now Brigantine city ; June 14, 1890 ) and Port Republic ( March 1, 1905 ).

After several failed attempts to incorporate in 1889, Wilbur was officially incorporated on August 11, 1890.

After a successful land auction in Harriman in 1890, the company established three subsidiaries: the East Tennessee Mining Company to administer the region's coal and iron extraction operations, the Harriman Coal & Iron Railroad Company to develop the local railroad system, and the Harriman Manufacturing Company to attract industries by providing start-up capital.

After visiting priesthood leaders in many settlements, President Woodruff left for San Francisco on September 3, 1890, to meet with prominent businessmen and politicians.

After his death, the king was buried in the church which he had built ; his original tomb has been lost, while his alleged remains are preserved in the shrine where he was reburied after being declared a saint ; his saintliness, however, was never very widely acknowledged outside the bishopric of Liège where he may still be venerated by tradition.

* 1858 – After the original Big Ben, a 14. 5 tonne bell for the Palace of Westminster had cracked during testing, it is recast into the current 13. 76 tonne bell by Whitechapel Bell Foundry.

After Ackermann's publication of his function ( which had three nonnegative integer arguments ), many authors modified it to suit various purposes, so that today " the Ackermann function " may refer to any of numerous variants of the original function.

After the original version of AMOS, Europress released two other versions: Easy AMOS, a simpler version for beginners, and AMOS Professional, a more advanced version with added features, such as a better IDE, ARexx support, a new UI sublanguage and new flow control constructs.

After a few divisions, the egg cell is placed into a uterus where it is allowed to develop into a fetus that is genetically identical to the donor of the original nucleus.

After Björkö came to be identified with ancient Birka, it has been assumed that the original name of Birka was simply Bierkø ( sometimes spelt Bjärkö ), an earlier form of Björkö.

After the downfall of Communism and the 1989 revolution, the original convention using î. Hr.

After Alston's death in 1977 a committee was formed, unable to raise funds for conservation on the original murals.

After completing his post-graduate work, he joined his father as assistant pastor of Boston's original North Church.

By the 18th century, map-makers started to give credit to the original engraver by printing the phrase " After original cartographer " on the work.

After executing it, he gave back the original French manuscript to Friedrich Schiller, from whom he had it.

After he evacuated from the island, he arranged the original music into a form suitable for musical performance.

After returning home, Thomas worked on Under Milk Wood in Wales, before sending the original manuscript to the BBC's Douglas Cleverdon on 15 October 1953.

After the end of the original series, Don sometimes produced additional " missing " chapters.

After some ten years of development, which Zamenhof spent translating literature into Esperanto as well as writing original prose and verse, the first book of Esperanto grammar was published in Warsaw in July 1887.

Director John Carpenter decided to add the original scenes into the special edition release as an extra only: " After we screened the rough cut, we realized that the movie didn't really start until Snake got to New York.

After several years of losing money, the original museum closed down in 1989 and the Evoluon was converted into a conference center, opening in 1998.

After Wood's departure following the band's debut record, Lynne wrote and arranged all of the group's original compositions and produced every album.

After several years of directing primarily lower-key live-action films, Hideaki Anno formed his own production studio and revisited his still-popular Evangelion franchise with the Rebuild of Evangelion tetralogy, a new series of films providing an alternate retelling of the original story.

After he became pope in 1978, John Paul II continued on the Catholic Theology of the Body of his predecessors with a series of lectures, entitled Theology of the Body, in which he talked about an original unity between man and women, purity of heart ( on the Sermon on the Mount ), marriage and celibacy and reflections on Humane Vitae, focusing largely on responsible parenthood and marital chastity.

After the lawsuit Hoshino Gakki abandoned the strategy of copying " classic " U. S. A. electric guitar designs — having already introduced a plethora of original designs.

After a few centuries it was abandoned for a second settlement, established in 6800 BCE, perhaps by an invading people who absorbed the original inhabitants into their dominant culture.

After German settlement, the original name of Cholberg evolved into.

After the defeat of Napoleon at Waterloo, many works seized by his armies were returned to their original owners.