Although many naturalists have studied aspects of animal behaviour throughout history, the modern discipline of ethology is generally considered to have begun during the 1930s with the work of Dutch biologist Nikolaas Tinbergen and Austrian biologists Konrad Lorenz and Karl von Frisch, joint winners of the 1973 Nobel Prize in Physiology or Medicine.

Although Hegel's philosophy of history is similar to Immanuel Kant's, and Karl Marx's theory of revolution towards the common good is partly based on Kant's view of history.

Although they were first discovered in 1883 by the German zoologist, Franz Eilhard Schulze ( 1840-1921 ) and since the 1970s more systematically analyzed by the German protozoologist, Karl Gottlieb Grell ( 1912-1994 ), a common name does not yet exist for the taxon ; the scientific name literally means " flat animals ".

Although he wanted Schiele to follow in his footsteps, and was distressed at his lack of interest in academia, he recognised Schiele's talent for drawing and unenthusiastically allowed him a tutor ; the artist Ludwig Karl Strauch.

Although O ' Rourke suggests that the first meta-analysis was a paper by Karl Pearson in the BMJ in 1904, in reality Pearson computed a simple average in this study and therefore it cannot be considered a meta-analysis.

Although Jacobi's elliptic functions are older and more directly relevant to applications, modern authors mostly follow Karl Weierstrass when presenting the elementary theory, because his functions are simpler, and any elliptic function can be expressed in terms of them.

Although non-specific concepts of madness have been around for several thousand years, the psychiatrist and philosopher Karl Jaspers was the first to define the three main criteria for a belief to be considered delusional in his 1913 book General Psychopathology.

Although Strauss was the most sought-after composer of dance music in the latter half of the 19th century, stiff competition was present in the form of Karl Michael Ziehrer and Émile Waldteufel ; the latter held a commanding position in Paris.

Although Karl Germer expelled Kenneth Grant from O. T. O.

Although he is credited for its discovery, Karl Wilhelm Feuerbach did not entirely discover the nine-point circle, but rather the six point circle, recognizing the significance of the midpoints of the three sides of the triangle and the feet of the altitudes of that triangle.

Although thoroughly " institutionalized ," Karl is deemed fit to be released into the outside world.

Although the problem arguably dates back to the Pyrrhonism of ancient philosophy, David Hume introduced it in the mid-18th century, with the most notable response provided by Karl Popper two centuries later.

* Although Karl Benz is credited with the modern automobile, which was created in 1885, it looked nothing like the automobile in the film, and vehicles like that would not be available until the 1920s.

Although Lassalle was a member of the Communist League, his politics were strongly opposed by Karl Marx and Friedrich Engels ; indeed Marx's essay Critique of the Gotha Program is written in part as a reaction to Lassalle's conception of the socialist state.

Although Sorge considered Friedrich Adolf Sorge, an associate of Karl Marx and Friedrich Engels, to be his grandfather, he was in fact his great-uncle.

Although György Lukács and Karl Korsch both attended the Arbeitswoche which had included a study of Korsch's Marxism and Philosophy, both were too committed to political activity and Party membership to join the Institut, although Korsch participated in publishing ventures for a number of years.

Although he was wary of the developing situation, the beginning of his active stance against the Nazis did not begin until the dismissal of Karl Planck, a well-known experimental physicist.

Although originally a Populist, after emigrating to Western Europe he established connections with the Social-Democratic movement of western Europe and began to study the works of Karl Marx and Friedrich Engels.

Although living in Bonn at the time of the paper's launch, Karl Marx seems to have been aware of the project from its inception and he began contributing articles to its pages, which drew notice from among the paper's readers.

Karl Rove stated in his 2010 memoirs that " Although Cleland personally blamed me for the ad, saying I was guilty of " character assassination " and responsible for " the biggest lie in America.

Although he had voted for war credits in August 1914, from July 1915 he opposed World War I and in 1917 he was among the founders of the USPD, which united anti-war socialists ( including reformists like Bernstein, " centrists " like Kautsky and revolutionary Marxists like Karl Liebknecht ).

Although initially an opponent of socialism, Bebel gradually was won over to socialist ideas through reading the pamphlets of Ferdinand Lassalle, which popularized the ideas of Karl Marx.

Although the orthodox Marxists in the party, led by Karl Kautsky, retained the Marxist theory of revolution as the official doctrine of the party, and it was repeatedly endorsed by SPD conferences, in practice the SPD leadership became increasingly reformist.

Although the orthodox Marxists in the party, led by Karl Kautsky, retained the Marxist theory of revolution as the official doctrine of the party, and it was repeatedly endorsed by SPD conferences, in practice the SPD leadership became more and more reformist.

Although both advocated workers ' rights, Henry George and Karl Marx were antagonists.

Although it began as a song cycle ( as reflected also by its title ), Arnold Schoenberg's Gurre-Lieder ( 1900 – 1903 / 1910 – 11 ) evolved into one of the century's largest secular cantatas.

Although Paul II was a committed opponent of humanist learning, he oversaw and approved the introduction of printing into the Papal States, first at Subiaco in 1464 by Arnold Pannartz and Konrad Sweynheim, and at Rome itself in 1467.

Although decommissioned by NASA in 2003, the NFAC is now being operated by the United States Air Force as a satellite facility of the Arnold Engineering Development Center ( AEDC ).

Although his mother Pauline, a native of Prague, was a piano teacher, Arnold was largely self-taught.

Although originally slated to premier with Hook, the SDDS project was delayed and instead premiered on June 17, 1993 with the Arnold Schwarzenegger film Last Action Hero.

Although Arnold had already been secretly negotiating with the British over a change of allegiance since May 1779, this news contributed to his decision to resign the command of Philadelphia.

Although Davis ' fundraising pace was criticized by his many detractors, Arnold Schwarzenegger would later collect contributions at a quicker rate during the early years of his governorship.

Although he retired in the late 1980s, he returned in 1993 in a minor supporting role in the Arnold Schwarzenegger film, Last Action Hero.

Although Robert Johnson died in 1938, James ( like many other musicians ) was strongly influenced by him, and by Kokomo Arnold and Tampa Red.

Although Montgomery was not permitted to launch an offensive on the scale he had wanted, George Marshall and Hap Arnold were eager to use the First Allied Airborne Army to cross the Rhine, so Eisenhower agreed to Operation Market-Garden.

Although he failed to achieve a first class degree in Greats, he won the Lothian and Arnold Prizes, the latter for an essay on Sir Thomas More ( about whom he confessed to having known almost nothing before commencing study, literally delivered as the clocks were chiming midnight on the day of the deadline ).

Although promoted to 1st lieutenant on April 10, 1913, Arnold was unhappy and requested a transfer to the Philippines.

Although Arnold was revered at the time, in retrospect Strachey sees his approach as very damaging.

Although the light infantry broke, General Fraser was trying to rally them, encouraging his men to hold their positions when Benedict Arnold arrived.

Although Helga has acknowledged her feelings regarding Arnold, she opts to keep them private by publicly denying and rejecting Arnold through displays of verbal abuse.

Although he defeated the American fleet in the Battle of Valcour Island and regained control of the lake, the rear guard defense managed by Benedict Arnold prevented further action to capture Ticonderoga or Crown Point in 1776.

Although, on the first ballot, Arnold Rüütel, a former leading communist and Chairman of the Presidium of the Supreme Council of the Estonian SSR, had led with 42 per cent of the total vote, the final choice for the nomination was made by Parliament, the Riigikogu, which was dominated by the Pro Patria Alliance.

Although trains and accessories of similar gauge and / or scale existed as early as 1927, modern commercially produced N scale models were first launched by the Arnold company of Nuremberg in 1962.

Although the original Arnold coupler has been joined by more functional and aesthetically pleasing designs ( see discussion below ), Arnold allowed use of the Rapido design by other manufacturers and so established a common standard to couple together rolling stock from different sources.

Although Johann Rudolf Meyer claimed to have reached the summit on an attempt in 1812 with guides Arnold Abbühl, Joseph Bortes and Aloys Volker, it is now accepted — after the research of John Percy Farrar in the Alpine Journal — that Jakob Leuthold and Johann Währen ( guides to Franz Joseph Hugi ) were the first to reach the summit via the north-west ridge on 10 August 1829 ( Hugi and others remained just below the top ).

Although he recorded a couple of songs with the band, he left the band before they recorded their first released single, " Arnold Layne " under the name The Pink Floyd.

Although Goehr's personal relationship to his father was not unproblematic, Walter Goehr had a determining influence on his son via his work as a conductor: the composers whose work Walter championed — Arnold Schoenberg, Claudio Monteverdi, Modest Mussorgsky, Olivier Messiaen — feature as a red-thread throughout Alexander's output.

Although, the fish-eating European White-tailed Eagles ( Haliaeetus albicilla ) could have caught fish in the River Trent, which lies a mere 4 miles south-east of Arnold, on the other side of the Mapperley Plains ridge.