Although single genes have little effect on complex traits and interact heavily both between themselves and with the external factors, research is underway to unravel possible molecular mechanisms underlying anxiety and comorbid conditions.

Although oxidation reactions are crucial for life, they can also be damaging ; plants and animals maintain complex systems of multiple types of antioxidants, such as glutathione, vitamin C, and vitamin E as well as enzymes such as catalase, superoxide dismutase and various peroxidases.

Although these encryption methods were more complex than previous schemes and required machines to encrypt and decrypt, other machines such as the British Bombe were invented to crack these encryption methods.

Although the words written in it are close enough to Italian and Latin for the non-Corsican speaker with a language background to follow, the pronunciation of those letters in English, French or Italian is not a guide to the pronunciation of Corsican, which follows complex rules.

Although most often used for matrices whose entries are real or complex numbers, the definition of the determinant only involves addition, subtraction and multiplication, and so it can be defined for square matrices with entries taken from any commutative ring.

Although the worship of images would eventually fall out of favour ( and be replaced by the iconoclastic fire temples ), the lasting legacy of the Achaemenids was a vast, complex hierarchy of Yazatas ( modern Zoroastrianism's Angels ) that were now not just evident in the religion, but firmly established, not least because the divinities received dedications in the Zoroastrian calendar, thus ensuring that they were frequently invoked.

Although the differences tend to be portrayed in polarized terms, both major coalitions have developed modified, nuanced and often complex positions.

Although production levels give an overview, the history of British cinema is complex, with various cultural movements developing independently.

Although many of his preparations today seem extravagant, he simplified and codified an even more complex cuisine that had existed beforehand.

Although this last approach required more complex mechanisms in many cases, it made for greater precision of results.

Although the Coalition was joined by other allies, the French Empire was also not alone since it now had a complex network of allies and submitted states.

Although the relationship between intellectual property and human rights is a complex one, there are moral arguments for intellectual property.

Although they accept that some complex systems and organs can be explained by evolution, they claim that organs and biological features which are irreducibly complex cannot be explained by current models, and that an intelligent designer must have created life or guided its evolution.

Although early Cretan coins occasionally exhibit multicursal patterns, the unicursal seven-course " Classical " design became associated with the Labyrinth on coins as early as 430 BC, and became widely used to represent the Labyrinth – even though both logic and literary descriptions make it clear that the Minotaur was trapped in a complex branching maze.

Although a complete theory of evolution also requires an account of how genetic variation arises in the first place ( such as by mutation and sexual reproduction ) and includes other evolutionary mechanisms ( such as genetic drift and gene flow ), natural selection appears to be the most important mechanism for creating complex adaptations in nature.

Although the nucleosome is a very stable protein-DNA complex, it is not static and has been shown to undergo a number of different structural re-arrangements including nucleosome sliding and DNA site exposure.

Although the simplest reflexes may be mediated by circuits lying entirely within the spinal cord, more complex responses rely on signal processing in the brain.

Although most of the country lies in the temperate belt, its climatic patterns are complex.

Although simple programs can be written in a few hours, programs that use complex mathematical formulas whose solutions can only be approximated or that draw data from many existing systems may require more than a year of work.

Although mathematically more complex, elliptic curves provide smaller key sizes and faster operations for approximately equivalent estimated security.

Although sections of the main Führerbunker were found, partially destroyed or filled in, another bunker complex was found further north that even the East German authorities had apparently missed, plus other cavities beneath land bordering the east side of Ebertstraße, although these turned out to be underground garages belonging to a former SS accommodation block.

Although constitutively expressed proteasomes can participate in this process, a specialized complex composed of proteins whose expression is induced by interferon gamma produces peptides of the optimal size and composition for MHC binding.

Although the oar can be conveniently thought of as a lever with a " fixed " pivot point in the water, the blade moves sideways and sternwards through the water, so that the magnitude of the propulsion force developed is the result of a complex interaction between unsteady fluid mechanics ( the water flow around the blade ) and solid mechanics and dynamics ( the handle force applied to the oar, the oar's inertia and bending characteristic, the acceleration of the boat and so on ).

Although the 709 was a superior machine to its predecessor, the 704, it was being built and sold at the time that transistor circuitry was supplanting vacuum tube circuits.

Although it appears it was never an official recommendation, by the mid-1950s the DRTE decided that the best way to really develop transistor techniques in a complex system was to build a computer.

Although one solution, commonly used at the time, was to build the core machinery out of tubes, for the DRTE machine this was considered one more challenge in transistor design.

Although vi was almost ubiquitous, he could not count on the local version working the way he expected.

Although a sizable force was routed near Bleiburg in the Bavarian March of Carinthia by Eberhard and the Count of Meran and another group was routed by Liutfried, count of Elsace, the Magyars repeatedly raided Germany.

Although about half of that money was spent on CPUs used in desktop or laptop personal computers, those count for only about 2 % of all CPUs sold.

Although detested by contemporaries, Tolstoy was made a count for his part in securing the throne for Catherine I.

Although The Planets remains Holst's most popular work, the composer himself did not count it among his best creations and later in life complained that its popularity had completely surpassed his other works.

Although the document of 1030 does not demonstrate that Amadeus and his father both held the rank of count simultaneously, Humbert's diploma of 1040 for the Diocese of Aosta was confirmed by his eldest son bearing the title count.

Although this category has a significant count, Japan is one of the few countries that reports it in the statistics as a separate category.

Although Taylor rose at the referee's count of six, he did not respond coherently to referee Richard Steele's questions and continued to hold onto the ropes all along.

Although there was some controversy in the final voting, for the first time, everyone was allowed to compete and have their votes count in the results.

Although Sancho was recognised as the ruler of Castile until his death, Ferdinand was granted the title " count " ( comes ) and was prepared to succeed in Castile.

Although the above is an argument for corporate taxation as opposed to the taxation of dividends, arguments for the taxation of income from capital would apply to both and on that count it can be argued that from a social policy standpoint it is unfair to tax income generated through active work at a higher rate than income generated through less active means ( although it might be said in defense that the ability to generate a material amount of dividend income can depend on years spent in active work pursuits ).

Although he began the race with a full war-chest and tied for first place with Dole in the 1995 Iowa Straw Poll, his campaign was fatally wounded when in an upset he lost the Louisiana Caucus on February 7, 1996 to Pat Buchanan ( the final delegate count was 13-8 ).

Although it is not mentioned, Musashi must have been well aware that this method would also be the most likely to have a higher death count than the others due to the sheer mass of enemies because more than one enemy could then attack a single soldier or trooper.

Although the collector could count the number of holes using a ruler, the usual practice is to use a perforation gauge, which has preprinted patterns of holes in a selection of common perforations, requiring one merely to line up the stamp's perforations with the closest match.

Although not himself a courtier, he was backed at court by Sosthene de la Rochefoucauld and Madame du Cayla, and in 1822 Louis XVIII gave him the title of count and made him formally prime minister.

Although it was not adopted by the Church, his method was essentially the same one known today as the Borda count, which is used in many academic institutions, competitions, and even some political jurisdictions, in original form and a number of variations.

Although Alexandre was unquestionably a product of Maria's affair with the Emperor, he was legalised by the old count Athenasius and thus bore the name of the counts of Colonna-Walewski.

Although his judges were biased against him, he was acquitted on each count, and the result was received enthusiastically by immense crowds inside and outside the court.

Although, following Theodoret, we have given the name Ophite to the system described by Irenaeus, it will have been seen that not only does the doctrine concerning the serpent form a very subordinate part of the system, but also that the place it assigns the serpent is very different from that given it by those whom we count as properly to be called Ophites.

Although he dominated the matches against the champion, Sammartino never won the title, winning most of his matches by count out or disqualification.

Although evidence is almost non-existent, it has been surmised that any vote was by count of head, but by the same token perhaps the fact so very little is said about actual votes suggests that most decisions, at least of a general kind, were reached more through persuasion and the weight by status of the county MPs.

Although the experimental section does not count towards the test-taker's score, it is unidentified and appears identical to the scored sections.

On September 25, 2006, Universal Press Syndicate president Lee Salem announced that the comic would not return, saying, " Although Aaron McGruder has made no statement about retiring or resuming The Boondocks for print newspapers ... newspapers should not count on it coming back in the foreseeable future.