However, it may have been for religious reasons, and would coincide with the development of religious practices thought to have occurred during the Upper Paleolithic .< ref > Nonetheless, it remains possible that Paleolithic societies never practiced cannibalism, and that the damage to recovered human bones was either the result of ritual post-mortem bone cleaning or predation by carnivores such as saber tooth cats, lions and hyenas.

However, it is the most conspicuous competition in grasslands, where, for example, cheetahs and hyenas are often killed by lion prides.

However, brown hyenas are aggressive scavengers, frequently appropriating the kills of black-backed jackals, cheetahs, and leopards, including adult male leopards.

However it is also possible that Middle Paleolithic cannibalism occurred for religious reasons which would coincide with the development of religious practices thought to have occurred during the Upper Paleolithic .< ref > Nonetheless it remains possible that Middle Paleolithic societies never practiced cannibalism and that the damage to recovered human bones was either the result of ritual post-mortem bone cleaning or predation by carnivores such as Saber tooth cats, lions and hyenas.

However, younger, weak or injured individuals may have been vulnerable to predation by tigers, pythons, crocodiles, Dinofelis, hyenas, bears, and Homo erectus.

However, claw-like nails are found in small-bodied callitrichids on all digits except the hallux or big toe, and in prosimians, which possess one or two laterally flattened toilet claws, used for grooming.

However one study of impalas found that impalas which were used by oxpeckers spent less time grooming themselves suggesting a reduction in parasite load.

However, they tend to emphasize strict " holiness standards " in dress, grooming and other areas of personal conduct that are not necessarily shared by other Pentecostal groups, at least not to the degree that is generally found in Oneness churches.

However, higher-ranking females give out and receive grooming the most.

However, Suffolk then made the mistake of attempting to undermine the rising power of Villiers by grooming another handsome young man to succeed him in James ' favor.

However after some time the chimpanzees used their own projected images for self-directed responding-such as grooming parts of the body before unseen without a mirror, picking their noses, making faces, and blowing bubbles at their own reflections.

However, as it ages, it acquires a sweet, earthy scent commonly likened to the fragrance of rubbing alcohol without the vaporous chemical astringency.

However, the Animal Welfare Act 2006 has made it illegal to remove their scent glands ( it is considered to be a cosmetic operation ), thus making them impractical as pets.

However, the boy found the scent of salt air much more alluring than the smell of shoe leather.

However, scent hounds are often stubborn, distracted, difficult to train, and some breeds will chase cats or other small mammals that resemble the prey they were bred to hunt.

However, Grenouille's ambitions are unmatched by technology: he cannot isolate the scent of inorganic materials, such as glass and iron, with the alembic that they use.

However, prior to his death, Rice exposed Kinney to the trigger scent ; X-23 goes into a murderous frenzy and kills her mother.

However thanks to Zander Rice lacing the trigger scent on her, Sarah was killed by Laura.

However, their scent and flavour is still vividly reminiscent of diethyl ether-presumably recreated via artificial means, in order to preserve the original flavour.

However, they more commonly communicate by scent marking and foot stamping.

However, he has two huge turnoffs to any prospective mates: his malodorous scent, and he cannot take ' no ' for an answer, blissfully convinced that the girl is flirting with him, even when she physically assaults him.

However, only a couple of months later a new problem arose: the company that commercially ran the circuit ( CENAV ), called in the receiver and went out of business, marking the end of " Circuit van Zandvoort ".

However, fifteen years later his views had changed, with Blench ( 2011 ) proposing instead that the noun-classifier system of Central Sudanic, commonly reflected in a tripartite general – singulative – plurative number system, triggered the development or elaboration of the noun-class system of the Atlantic – Congo languages, with tripartite number marking surviving in the Plateau and Gur languages of Niger – Congo, and the lexical similarities being due to loans.

However, it is customary for synagogues to conduct a siyum ( ceremony marking the completion of a section of Torah learning ) right after morning prayers, and the celebratory meal that follows cancels the firstborn's obligation to fast.

However, Rush's triple CD live album, Rush in Rio ( 2003 ), was certified gold by the RIAA, marking the fourth decade in which a Rush album had been released and certified at least gold.

However, they exhibit anatomical differences marking them off as a group distinct from their Carboniferous and Recent relatives.

However, instead of marking moves on a piece of paper, the games is played verbally, with each player calling out which locations they take.

However the emperor permanently barred Li Ying and his associates from serving in office, marking the beginning of the Partisan Prohibitions.

However, in 1529 a lawyer, Sir Thomas More, was appointed as Chancellor, marking the beginning of a new era.

However, his popularity suffered from the economic downturn that followed the 1973 energy crisis, marking the end of the " thirty glorious years " after World War II, combined with the official discourse that the " end of the tunnel was near ".

However, by marking the overlap range from multiple balances, one can achieve a considerable level of accuracy.

However, the first real public courts have recently opened in the Westway sports centre in London's White City, marking a possible change in fortunes for Eton Fives as a minor sport.

However this event does signify the end of the Danish-ruled Kalmar Union, so in a sense it is a marking of Swedish independence, though the event occurred so long ago that it does not have as strong of a presence in the social consciousness as does, for example, Norway's Syttende Mai ( 17 May ).

However, the method was prone to supermarkets cheating by temporarily marking down shelf prices while the surveyor was in store, and the method was scrapped in 2003 after the Dunedin Pak ' nSave was caught in the act, marking down prices by over half in some cases.

However, a forward is not traded on an exchange and thus does not have the interim partial payments due to marking to market.

However, it is believed that black widow young have at least some sort of marking on their abdomens.

However, such an arrangement did not materialize, and in October 2007, Pride Worldwide's Japanese staff was laid off, marking the end of the organization as an active fight promoter.

However, the smaller structures may be built just as signs, marking the mountain paths or boundaries of owned land.

However, this change in the competition format did not eliminate the need for marking based on hand-written programs.

However, Rough Trade's own 1983 album Weapons failed to place a single on the Canadian charts, marking the beginning of the group's commercial decline.

However, he leaves behind a part of a map marking the location of Totenkopf's base.

However, it was later realised that the multiplicity of urban route marking schemes led to confusion.

However, they failed to win the Grey Cup, marking the 2000s as the first decade since the 1890s that Hamilton failed to win a national championship.

However not all prepositions trigger prepositional case marking, and a small group of prepositions which are termed compound mark their objects with genitive case, these prepositions being historically derived from the fusion of a preposition plus a following noun which has become grammaticalised.