Back on Chapterhouse, Odrade confronts Duncan and forces him to admit that he is a Mentat, proving that he retains the memories of his many ghola lives.

Odrade is a loose cannon ; she does not obey normal Bene Gesserit prohibitions about love, and is also Teg's biological daughter.

This agreement causes consternation among the Bene Gesserit, but Odrade realizes that Taraza's plan is to destroy Rakis.

The new Mother Superior Darwi Odrade recognizes that the threat of this unknown enemy is greater than that of the Honored Matres, and forms another bold plan.

She immediately succeeds Odrade as Mother Superior of the Bene Gesserit, joining the two forces under a single leader in an uneasy truce that is hoped will be able to defeat the unknown enemy.

In Chapterhouse Dune, Darwi Odrade is Mother Superior, a contentious choice ratified by the fact that she was present at the previous Mother Superior's death, and has her in Other Memory ; she represents the most continuous line of leadership.

In Heretics of Dune ( 1984 ), one descendant of Siona Atreides is Miles Teg, renowned commander of the military forces of the Bene Gesserit, who bears a remarkable resemblance to his ancestor, Duke Leto I. Teg's secret daughter is Darwi Odrade ( a variant on Atreides ), a Bene Gesserit sister who eventually becomes Mother Superior of the Sisterhood during their ongoing struggle with the fierce Honored Matres.

In Heretics of Dune, the Mother Superior of the Bene Gesserit Taraza ( and her immediate successor, Odrade ) manage to save one sandworm from Rakis before it is destroyed by the Honored Matres.

Taraza is killed ; her successor Darwi Odrade takes the worm to Chapterhouse.

Eventually, the core of the Honored Matres is destroyed according to the plan of the Bene Gesserit leader, Mother Superior Darwi Odrade.

Bene Gesserit leader Odrade is surprised, but she and Miles Teg soon realize they have fallen into a trap.

Bene Gesserit leader Darwi Odrade notes that " Scytale admits to memories of Muad ' Dib's times ," and Herbert clarifies that Scytale is technically a clone, as the cells used to resurrect him had been taken from his living predecessor rather than a corpse.

During the events of Heretics of Dune, it is revealed that Teg had fathered other children during his younger years, and he discovers that Reverend Mother Darwi Odrade is one of them.

At the beginning of Chapterhouse: Dune ( 1985 ), a ghola of Teg is birthed on orders from his daughter, Odrade, who is now Mother Superior of the Bene Gesserit after Taraza's death at the end of Heretics of Dune.

Murbella, a captive Honored Matre indoctrinated into the Bene Gesserit, kills the Honored Matre leader Logno as Bene Gesserit Mother Superior Odrade is killed, and Murbella manages to secure the leadership of both groups.

Odrade relaxes the strictures, and for the first time ever, a man – Duncan – is allowed to watch a spice ceremony.

Odrade watches with equal trepidation because Murbella is vital to her plan to deal with the Honored Matres and to reform the Sisterhood.

Murbella, Odrade is pleased to find, takes and survives the hard path to becoming a Reverend Mother.

Odrade is killed in the melee.

Taraza also sends Reverend Mother Darwi Odrade to take command of the Bene Gesserit keep on Rakis.

Odrade permits Duncan to watch Murbella undergo the spice agony, making him the first man ever to do so.

Odrade, however, had planned for the possible failure of the Bene Gesserit attack and left Murbella instructions for a last desperate gamble.

At the Battle of Junction, the Honored Matre stronghold, both Odrade and Great Honored Matre Logno perish, giving Murbella, a former Honored Matre assimilated into the Bene Gesserit, the chance to ascend to both positions, carrying out the rest of Odrade's plan by unifying the orders.

However in Chapterhouse: Dune both Odrade and Murbella comment that these skills are nowhere near as potent as the Bene Gesserit Voice.

Odrade, however, had planned for the possible failure of the Bene Gesserit attack and left Murbella instructions for a last desperate gamble.

Murbella goes through a manic phase after becoming a Reverend Mother, and her feelings for Duncan are not the same – a fact that worries Odrade as much as Duncan.

The potential failure of the Bene Gesserit attack was planned for by Odrade, who left Murbella instructions for a last desperate gamble.

Murbella shares with Odrade and takes charge of the Honored Matres, who are awed by her physical prowess.

He tries to assassinate Taraza but Odrade convinces him that the Sisterhood shares the religious beliefs of the Bene Tleilax.

Odrade continues molding Scytale, with Sheeana showing him a baby sandworm, the Bene Gesserit's own long term supply of spice, and destroying Scytale's main bargaining card.

Odrade becomes temporary leader of the Bene Gesserit before escaping with Sheeana into the desert on a worm.

We are presented with the greatest face of this in Mother Superior Darwi Odrade, a female descendant of Siona Atreides and Duncan Idaho.

He takes the ship to Rakis to meet up with Bene Gesserits Sheeana and Odrade.

Now in command of the Bene Gesserit, Mother Superior Darwi Odrade continues to develop her drastic, secret plan to overcome the Honored Matres.

Odrade next calls a meeting of all the Bene Gesserit, announcing her plan to attack the Honored Matres.

Odrade calls a Convocation ( a meeting of all the Bene Gesserit ) announcing her plan to attack the Honored Matres.

Odrade believes that the Bene Gesserit made a mistake in fearing emotion, and that in order to evolve, the Bene Gesserit must learn to accept emotions.

Odrade then confronts Sheeana, discovering that Duncan and Sheeana have been allied together for some time.

Sheeana does not reveal that they have been considering the option of reawakening Teg's memory through Imprinting, nor does Odrade discover that Sheeana has the keys to Duncan's no-ship prison.

Odrade appoints him again as Bashar of the military forces of the Sisterhood for the assault on the Honored Matres.

Odrade then goes to meet the Great Honored Matre.

Her first act surprises Odrade greatly.

Too late Odrade and Teg realize they have fallen into a trap, and the Honored Matres use a mysterious weapon to turn defeat into victory, as well as capturing Odrade.

Odrade starts training Sheeana as a Bene Gesserit.

She and Odrade meet Waff again on Rakis.

When they arrive, Teg finds Odrade and Sheeana and their giant worm.